1. Evolutionary Genomics Group; Department of Botany and Zoology; Stellenbosch University; Private Bag X1 Matieland 7602 South Africa
2. Centre for Invasion Biology; Department of Zoology; University of Johannesburg; PO Box 524 Auckland Park South Africa
3. Department of Conservation Ecology and Entomology; Stellenbosch University; Private Bag X1 Matieland 7602 South Africa