1. Department of Biological Chemistry and Crop Protection; Rothamsted Research; Harpenden Hertfordshire AL5 2JQ UK
2. Department of Plant Biology and Crop Science; Rothamsted Research; Harpenden Hertfordshire AL5 2JQ UK
3. Institute of Life Science; College of Medicine; Swansea University; Swansea SA2 8PP UK
4. Bayer CropScience AG; Research Disease Control; 40789 Monheim Germany
5. School of Agriculture, Policy and Development; University of Reading; Whiteknights Reading RG6 6AS UK