1. Graduate School of Natural Science, Okayama University, Okayama 700‐8530, Japan
2. Faculty of Agriculture, Okayama University, Okayama 700‐8530, Japan
3. Faculty of Agriculture, Yamagata University, Tsuruoka, Yamagata 997‐8555, Japan
4. Graduate School of Bioagricultural Sciences, Nagoya University, Nagoya, Aichi 464‐8601, Japan