1. Pohang University of Science and Technology–University of Zurich Cooperative Laboratory, Department of Integrative Bioscience and Biotechnology, World Class University Program, Pohang University of Science and Technology, Pohang 790‐784, Korea
2. Advanced Converging Technology Research Laboratory, Department of Chemical and Biomolecular Engineering, Yonsei University, Seoul 120‐749, Korea
3. Plant and Soil Science Laboratory, Department of Agriculture and Ecology, Faculty of Life Sciences, University of Copenhagen, Thorvaldsensvej 40, DK‐1871 Frederiksberg C, Denmark
4. Institute of Plant Biology, University of Zurich, CH‐8008 Zurich, Switzerland