1. State Key Laboratory of Biocontrol Guangdong Key Lab of Plant Resources, School of Life Sciences, Southern Marine Science and Engineering Guangdong Laboratory (Zhuhai), Sun Yat‐Sen University 510275 Guangzhou Guangdong China
2. Hainan Academy of Forestry (Hainan Academy of Mangrove) 571199 Haikou Hainan China
3. Department of Botany and Biodiversity Research Centre University of British Columbia Vancouver British Columbia Canada