1. Environment Centre Wales Bangor University Gwynedd LL57 2UW UK
2. Centre for Microscopy Characterisation and Analysis University of Western Australia Crawley WA 6009 Australia
3. School of Agriculture, Food and Rural Development Newcastle University Newcastle upon Tyne NE1 7RU UK
4. Soil Biology and Molecular Ecology Group School of Earth and Environment Institute of Agriculture University of Western Australia Crawley WA 6009 Australia