1. Institute of Evolution and Ecology, School of Life Sciences Central China Normal University Wuhan 430079 China
2. Key Laboratory of State Forestry Administration on Biodiversity Conservation in Karst Mountainous Areas of Southwestern China, School of Life Sciences Guizhou Normal University Guiyang 550025 China
3. State Key Laboratory of Systematic and Evolutionary Botany, Institute of Botany Chinese Academy of Sciences Beijing 100093 China