1. State Key Laboratory of Tree Genetics and Breeding, Key Laboratory of Silviculture of the State Forestry Administration, Research Institute of Forestry Chinese Academy of Forestry Beijing 100091 China
2. Collaborative Innovation Center of Sustainable, Forestry in Southern China Nanjing Forestry University Nanjing 210037 China
3. Department of Plant Biology, Linnean Centre for Plant Biology Swedish University of Agricultural Sciences Uppsala 75007 Sweden
4. Forestry Research Institute of Altay Region, Altay Xinjiang 836500 China
5. Shenzhen Branch, Guangdong Laboratory for Lingnan Modern Agriculture, Genome Analysis Laboratory of the Ministry of Agriculture, Agricultural Genomics Institute at Shenzhen Chinese Academy of Agricultural Sciences Shenzhen 518120 China