1. Instituto Pirenaico de Ecología (IPE-CSIC); Avda Nuestra Señora de la Victoria 16 22700 Jaca Spain
2. Department of Biodiversity and Evolutionary Biology; Museo Nacional de Ciencias Naturales (MNCN-CSIC); C/José Gutiérrez Abascal 2 28006 Madrid Spain
3. Department of Ecology and Evolution; University of Lausanne; Biophore 1015 Lausanne Switzerland
4. Fundación Araid; Edificio CEEI Aragón; C/María de Luna 11 50018 Zaragoza Spain