1. Department of Biochemistry, Genetics and Microbiology, Forestry and Agricultural Biotechnology Institute (FABI); University of Pretoria; Pretoria 0002 South Africa
2. Department of Conservation Ecology and Entomology; Stellenbosch University, Private ; Bag X1 Matieland 7602 South Africa
3. Department of Botany and Zoology; Stellenbosch University, Private; Bag X1 Matieland 7602 South Africa