1. Faculty of Science; Leiden University; P.O. Box 9502 2300 RA Leiden The Netherlands
2. Biodiversity Dynamics Research Group; Naturalis Biodiversity Center; Vondellaan 55, P.O. Box 9517 2300 RA Leiden The Netherlands
3. Institute for Biodiversity and Ecosystem Dynamics (IBED), Computational Geo-Ecology; University of Amsterdam; Science Park 904 1098 HX Amsterdam The Netherlands
4. Instituto Multidisciplinario de Biología Vegetal (IMBIV); CONICET; Universidad Nacional de Córdoba; CC 495 5000 Córdoba Argentina
5. Fundación ProYungas; Perú 1180, 4107 Yerba Buena Tucumán Argentina