1. NIMBioS University of Tennessee Knoxville Tennessee USA
2. CEFE, Univ Montpellier, CNRS, EPHE IRD Montpellier France
3. Department of Forestry Wildlife and Fisheries, University of Tennessee Knoxville Tennessee USA
4. Department of Ecology and Evolutionary Biology University of Tennessee Knoxville Tennessee USA
5. Department of Mathematics University of Tennessee Knoxville Tennessee USA