1. Research Center for Micro-Nano Technology; University of Hosei; Midori-cho 3-11-15 Koganei Tokyo 184-0003 Japan
2. Section of Cell Biology and Pathology; Department of Alzheimer's Disease Research; Center for Development of Advanced Medicine for Dementia; National Center for Geriatrics and Gerontology; Morioka 7-430 Obu Aichi 474-8511 Japan
3. Department of Biophysics and Biochemistry; Graduate School of Science; The University of Tokyo; Yayoi 2-11-16 Bunkyo-ku Tokyo 113-0032 Japan
4. Department of Molecular and Developmental Biology; Institute of Medical Science; The University of Tokyo; Shirokane-dai 4-6-1 Minatoku-ku Tokyo 108-8639 Japan