1. Qingdao Institute of Marine GeologyChina Geological Survey Qingdao 266071 China
2. Laboratory for Marine Mineral ResourcesQingdao National Laboratory for Marine Science and Technology Qingdao 266071 China
3. Laboratory for Marine GeologyQingdao National Laboratory for Marine Science and Technology Qingdao 266071 China
4. Department of BiologyUniversity of Bergen Bergen 5808 Norway