1. Palaeontological Center Bohai University Jinzhou 121013 Liaoning China
2. Shenzhen Key Laboratory for Orchid Conservation and Utilization National Orchid Conservation Center of China and Orchid Conservation & Research Center of Shenzhen Shenzhen 518114 Guangdong China
3. State Key Laboratory of Palaeobiology and Stratigraphy Nanjing Institute of Geology and Palaeontology, CAS Nanjing 210008 China
4. Key Laboratory of Tropical Forest Ecology Xishuangbanna Tropical Botanical Garden, CAS Mengla 666303 Yunnan China