1. School of Life Sciences; University of Nottingham Medical School; Nottingham NG7 2UH UK
2. Neuroscience Division, Medical Education Institute; Ninewells Hospital and Medical School; University of Dundee Dundee, DD1 9SY UK
3. School of Physiology, Pharmacology and Neuroscience; University of Bristol; Bristol BS8 1TD UK
4. Centre for Integrative Physiology; University of Edinburgh; Edinburgh EH8 9XD UK