1. College of Agriculture & Biotechnology; Zhejiang University; Zijingang Campus; Hangzhou 310058 People's Republic of China
2. Zhejiang Provincial Key Laboratory of Horticultural Plant Integrative Biology; Zhejiang University; Zijingang Campus; Hangzhou 310058 People's Republic of China
3. The State Agriculture Ministry Laboratory of Horticultural Plant Growth; Development and Quality Improvement; Zhejiang University; Zijingang Campus; Hangzhou 310058 People's Republic of China
4. New Zealand Institute for Plant & Food Research Limited; Private Bag 92169 Auckland New Zealand
5. Boyce Thompson Institute for Plant Research; Ithaca NY 14853 USA
6. US Department of Agriculture/Agriculture Research Service; Robert W. Holley Centre for Agriculture and Health; Ithaca NY 14853 USA