1. Graduate School of Systems Life Sciences Kyushu University Graduate School Nishi‐ku Fukuoka 819‐0395 Japan
2. Department of Biology, Faculty of Sciences Kyushu University Nishi‐ku Fukuoka 819‐0395 Japan
3. Present address: Department of Infectious Disease Control, International Research Center for Infectious Diseases, The Institute of Medical Science The University of Tokyo Tokyo 108‐8639 Japan
4. Medical Institute of Bioregulation Kyushu University Higashi‐ku Fukuoka 812‐8582 Japan