1. RIKEN Center for Sustainable Resource Science (CSRS); 3-1-1 Koyadai Tsukuba Ibaraki 305-0074 Japan
2. Japan International Research Center for Agricultural Sciences (JIRCAS); 1-1 Ohwashi Tsukuba Ibaraki 305-8686 Japan
3. RIKEN Plant Science Center; 1-7-22 Suehiro-cho Tsurumi-ku Yokohama, Kanagawa 230-0045 Japan
4. RIKEN Center for Sustainable Resource Science (CSRS); 2-1 Hirosawa Wako Saitama 351-0198 Japan
5. Graduate School of Agricultural and Life Sciences; The University of Tokyo; Bunkyo-ku Tokyo Japan