1. Department of Ecology and Evolutionary Biology; Cornell University; Corson Hall Ithaca NY 14853-2701 USA
2. Computational and Molecular Population Genetics (CMPG); Institute of Ecology and Evolution; University of Bern; Baltzerstrasse 6 CH-3012 Bern Switzerland
3. School of Biosciences; Cardiff University; Sir Martin Evans Building Museum Avenue Cardiff CF10 3AX UK
4. Swiss Institute of Bioinformatics; Genopode; CH 1015 Lausanne Switzerland