1. Department of Soil Science of Temperate Ecosystems; Georg-August University of Göttingen; Büsgenweg 2 37077 Göttingen Germany
2. Department of Agricultural Soil Sciences; Georg-August University of Göttingen; Büsgenweg 2 37077 Göttingen Germany
3. Sustainable Soils and Grassland Systems Department; Rothamsted Research; North Wyke Okehampton EX20 2SB UK
4. UCD School of Agriculture and Food Science, Agriculture and Food Science Centre; University College Dublin; Belfield, Dublin 4 Ireland