1. Department of Ecology and Evolution; University of Lausanne; 1015 Lausanne Switzerland
2. Swiss Institute of Bioinformatics; 1015 Lausanne Switzerland
3. Vital-IT; Swiss Institute of Bioinformatics; 1015 Lausanne Switzerland
4. Department of Evolutionary Biology; Evolutionary Biology Centre; Uppsala University; Norbyvägen 18D 75236 Uppsala Sweden