1. Department of Oceanography; School of Ocean and Earth Science and Technology; University of Hawai′i at Mānoa; Honolulu HI 96822 USA
2. School of Biological and Biomedical Sciences; Durham University; South Road Durham DH1 3LE UK
3. Hawai′i Institute of Marine Biology; University of Hawai′i at Mānoa; Kāne′ohe HI 96744 USA
4. Department of Ichthyology; California Academy of Sciences; Golden Gate Park 55 Music Concourse Drive San Francisco CA 94118 USA