1. Department of Ecology and Evolution; University of Lausanne; 1015 Lausanne Switzerland
2. Swiss Institute of Bioinformatics; Quartier Sorge 1015 Lausanne Swizterland
3. Landscape Dynamics; Swiss Federal Research Institute WSL; 8903 Birmensdorf Switzerland
4. Department of Plant Biology and Ecology; University of the Basque Country UPV/EHU; 48080 Bilbao, Spain and IKERBASQUE, Basque Foundation for Science 48011 Bilbao Spain