1. Philip Leverhulme Equine Hospital; Institute of Veterinary Science University of Liverpool; Chester High Road Neston CH64 7TE UK
2. Section of Veterinary Pathology; Institute of Veterinary Science University of Liverpool; Chester High Road Neston CH64 7TE UK
3. Institute of Ageing and Chronic Disease University of Liverpool; 6 West Derby Street Liverpool L7 8TX UK