1. Borneo Marine Research Institute; Universiti Malaysia Sabah; Kota Kinabalu Sabah Malaysia
2. Department of Fisheries; Graduate School of Agriculture; Kinki University; Nara Japan
3. Department of Aqua Bioscience and Industry; Faculty of Bioindustry; Tokyo University of Agriculture; Hokkaido Japan
4. Fisheries Laboratory; Kinki University; Shirahama Wakayama Japan
5. Fisheries Laboratory; Kinki University; Uragami Wakayama Japan