1. Department of Invertebrates; Natural History Museum of Bern; Bernastrasse 15 3005 Bern Switzerland
2. Department of Entomology; Natural History Museum of Denmark; Zoological Museum; University of Copenhagen; Universitetsparken 15 2100 Copenhagen Denmark
3. Center of Macroecology, Evolution and Climate; Biological Institute; Universitetsparken 15 2100 Copenhagen Denmark