1. Department of Marine Ecology; Institut Mediterrani d'Estudis Avançats; IMEDEA (CSIC-UIB); C/Miquel Marquès 21 07190 Esporles, Balearic Islands Spain
2. Department of Marine Sciences and Applied Biology; University of Alicante; P. O. Box 99 03080 Alicante Spain
3. Department of Biology and Ecology of Fishes; Leibniz-Institute of Freshwater Ecology and Inland Fisheries; Müggelseedamm 310 12587 Berlin Germany