1. Graduate School of Bioresource & Bioenvironmental Sciences; Kyushu University; Fukuoka Japan
2. Kyushu Regional Breeding Office; Forest Tree Breeding Center, Forestry and Forest Products Research Institute; Koshi Japan
3. Forest Tree Breeding Center; Forestry and Forest Products Research Institute; Hitachi Japan
4. Faculty of Agriculture; Kyushu University; Fukuoka Japan