1. Department of Cell Science; Research Institute for Frontier Medicine, Sapporo Medical University School of Medicine; Sapporo Japan
2. Sapporo Medical University School of Medicine; Sapporo Japan
3. Department of Otolaryngology; Sapporo Medical University School of Medicine; Sapporo Japan
4. Department of Human Immunology; Research Institute for Frontier Medicine, Sapporo Medical University School of Medicine; Sapporo Japan