1. Department of Agriculture and Fisheries; Robert Wicks Pest Animal Research Centre; Biosecurity Queensland; Toowoomba Queensland 4350 Australia
2. University of Southern Queensland; Institute for Agriculture and the Environment; Toowoomba Queensland 4350 Australia
3. Department of National Parks, Sport and Racing; Queensland Parks and Wildlife Service; Fraser Island Queensland Australia
4. School of Agriculture and Food Sciences; The University of Queensland; Gatton Queensland Australia
5. Department of Agriculture and Fisheries; Biosecurity Queensland; Southport Queensland Australia
6. Department of Agriculture and Fisheries; Animal Biosecurity and Welfare; Biosecurity Queensland; Coopers Plains Queensland Australia