1. Centre of Excellence in Biological Interactions, Division of Ecology, Evolution & Genetics, Research School of Biology; Australian National University; Canberra ACT 2601 Australia
2. Institute of Integrative Biology; University of Liverpool; Liverpool L69 7ZB United Kingdom
3. Biosciences, University of Exeter; Cornwall Campus; Penryn TR10 9FE United Kingdom
4. Institute of Evolutionary Biology and Environmental Sciences; University of Zurich; Winterthurerstrasse 190 CH-8057 Zurich Switzerland