1. Key Laboratory for Plant Diversity and Biogeography of East Asia; Kunming Institute of Botany, Chinese Academy of Sciences; Kunming Yunnan 650201 China
2. Plant Germplasm and Genomics Center, Germplasm Bank of Wild Species; Kunming Institute of Botany, Chinese Academy of Sciences; Kunming Yunnan 650201 China
3. Kunming College of Life Sciences; University of Chinese Academy of Sciences; Kunming Yunnan 650201 China
4. Institute of Plant Science; School of Life Science, Yunnan University; Kunming Yunnan 650091 China
5. Department of Botany; California Academy of Sciences; 55 Music Concourse Drive San Francisco CA 94118 USA