1. Key Laboratory of Tropical Forest Ecology Xishuangbanna Tropical Botanical Garden Chinese Academy of Sciences Mengla Yunnan 666303 China
2. University of Chinese Academy of Sciences Beijing 100049 China
3. State Key Laboratory for Conservation and Utilization of Subtropical Agro‐bioresources College of Forestry Guangxi University Nanning Guangxi 530004 China
4. University of Wisconsin‐Milwaukee Milwaukee PO Box 413 WI 53201 USA
5. Smithsonian Tropical Research Institute Apartado 2072 Balboa Panama
6. Forest Ecology and Forest Management Group Wageningen University PO Box 476700 AA Wageningen the Netherlands