1. Department of Genetics, Evolution & Environment; University College London; Darwin Building Gower Street London WC1E 6BT UK
2. Department of Earth Sciences; University College London; Gower Street London WC1E 6BT UK
3. Department of Earth Science and Engineering; Imperial College London; South Kensington Campus London SW7 2AZ UK
4. School of Earth Sciences; University of Bristol; Wills Memorial Building Queens Road Clifton BS8 1RJ UK