1. Hainan Key Laboratory for Sustainable Utilisation of Tropical Bioresource, Institute of Tropical Agriculture and ForestryHainan University Haikou China
2. National Key Laboratory of Crop Genetic Improvement and National Center of Plant Gene Research (Wuhan)Huazhong Agricultural University Wuhan China
3. College of Life Science and TechnologyHuazhong Agricultural University Wuhan China
4. Institute of Biological, Environmental and Rural SciencesAberystwyth University Aberystwyth UK