1. Royal (Dick) School of Veterinary Studies, University of Edinburgh, Summerhall, Edinburgh EH9 1QH
2. *Department of Pathology and Microbiology, University of Bristol, Langford House, Langford, Bristol BS40 5DU
3. †Department of Veterinary Clinical Science and Animal Husbandry, University of Liverpool, Leahurst, Neston, South Wirral L64 7TE