1. Section for Evolutionary Ecology Department of Biology Lund University Sölvegatan 37 SE‐223 62 Lund Sweden
2. Scottish Centre for Ecology and the Natural Environment Institute of Biodiversity, Animal Health and Comparative Medicine University of Glasgow Rowardennan G63 0AW UK
3. Department of Forestry & Environmental Resources North Carolina State University 11 W. Jones Street Raleigh NC 27601 USA