1. Research Institute for Microbial Diseases Osaka University Suita Osaka Japan
2. Graduate School of Pharmaceutical Sciences Osaka University Suita Osaka Japan
3. Priority Organization for Innovation and Excellence Kumamoto University Kumamoto Japan
4. Division of Reproductive Biology, Institute of Resource Development and Analysis Kumamoto University Kumamoto Japan
5. Center for Drug Discovery and Department of Pathology & Immunology Baylor College of Medicine Houston Texas USA
6. The Institute of Medical Science The University of Tokyo Minato‐ku Tokyo Japan