1. School of Natural Resources and the Environment University of Arizona College of Agriculture and Life Sciences 1064 E Lowell Street Tucson AZ 85721‐0001 U.S.A.
2. Department of Environmental Science University of Arizona 1177 E 4th St Tucson AZ 85721‐0001 U.S.A.
3. College of Science and Engineering Flinders University Adelaide South Australia 5001 Australia