1. Budongo Conservation Field Station; P.O. Box 362 Masindi Uganda
2. Department of Forestry, Biodiversity and Tourism; Makerere University; P.O. Box 7062 Kampala Uganda
3. Wildlife Conservation Society; 2300 Southern Boulevard Bronx NY 10460 USA
4. Conservation Science Group; Department of Zoology; Cambridge University; Pembroke Rd Cambridge UK
5. Department of Biology; University of York; Wentworth Way York YO10 5DD UK
6. School of Biological Sciences; The University of Queensland; St Lucia Brisbane Qld 4072 Australia
7. Department of Biological Anthropology; University of Oxford; Banbury Rd Oxford UK