1. Biodiversity Program, Taiwan International Graduate Program, Academia Sinica National Taiwan Normal University Taipei 115201 Taiwan
2. Biodiversity Research Center Academia Sinica Taipei 115201 Taiwan
3. Department of Life Science National Taiwan Normal University Taipei 106209 Taiwan
4. Native Plant Centre, Horticulture & Community Gardening Division National Parks Board 1 Cluny Road Singapore 259569 Singapore
5. Department of Biological Sciences, Graduate School of Science The University of Tokyo 2‐11‐16 Yayoi, Bunkyo‐ku Tokyo 113‐0032 Japan
6. Institut de Systématique, Évolution, Biodiversité, Muséum national d'Histoire naturelle, Sorbonne Université, Ecole Pratique des Hautes Etudes, CNRS Université des Antilles Paris 75005 France
7. Department of Plant Biology University of Vermont Burlington VT 05405 USA
8. Museum of New Zealand Te Papa Tongarewa Wellington 6011 New Zealand
9. Taiwan Forestry Research Institute Taipei 100060 Taiwan