1. Department of Neuroscience, Okayama University Graduate School of Medicine; Dentistry and Pharmaceutical Sciences; Kita-ku Okayama 700-8558 Japan
2. CREST; Japan Science and Technology Agency; Kita-ku Okayama 700-8558 Japan
3. Department of Cell Physiology, Okayama University Graduate School of Medicine; Dentistry and Pharmaceutical Sciences; Kita-ku Okayama 700-8558 Japan
4. Department of Molecular Physiology, Faculty of Life Sciences; Kumamoto University; Kumamoto 860-8556 Japan
5. Department of Urology, Okayama University Graduate School of Medicine; Dentistry and Pharmaceutical Sciences; Kita-ku Okayama 700-8558 Japan