1. Key Laboratory of Freshwater Fisheries and Germplasm Resources Utilization, Ministry of Agriculture, Freshwater Fisheries Research Center; Chinese Academy of Fishery Sciences; No. 9 Shanshui East Road Wuxi 214081 China
2. Department of Aquaculture; Wuxi Fisheries College, Nanjing Agriculture University; Wuxi 214081 China
3. Department of Forestry and Natural Resources; Purdue University; West Lafayette Indiana 47907 USA
4. Yancheng Institute of Technology; Yancheng 224051 China