1. Department of Animal Ecology and Tropical Biology, Biocenter; University of Würzburg; Am Hubland D-97074 Würzburg Germany
2. Department of Evolutionary Biology and Animal Ecology; University of Freiburg; Hauptstrasse 1 D-79114 Freiburg Germany
3. Institute of Environmental Sciences (ICAM); University of Castilla-La Mancha; Avenida Carlos III, s/n; E-45071 Toledo Spain
4. Center for Computational and Theoretical Biology, Campus Nord; University of Würzburg; Am Hubland D-97074 Würzburg Germany
5. Department of Animal Ecology I, Bayreuth Center for Ecology and Environmental Research (BayCEER); University of Bayreuth; D-95440 Bayreuth Germany