1. Guangxi Key Laboratory of Buffalo Genetics, Reproduction and Breeding Guangxi Buffalo Research Institute, Chinese Academy of Agricultural Sciences Nanning China
2. State Key Laboratory for Conservation and Utilization of Subtropical Agro‐Bioresources College of Animal Science and Technology, Guangxi University Nanning Guangxi China
3. Key Laboratory of Buffalo Genetics, Breeding and Reproduction Technology Ministry of Agriculture and Rural Affairs Nanning China
4. College of Animal Science and Technology Guangxi Vocational University of Agriculture Nanning China