1. Key laboratory of Horticultural Plant Biology, Ministry of Education Huazhong Agricultural University Wuhan China
2. College of Plant Science and Technology Huazhong Agricultural University Wuhan China
3. Flower Research Institute of Yunnan Academy of Agricultural Sciences National Engineering Research Center For Ornamental Horticulture Kunming China
4. Guangdong Key Laboratory of Ornamental Plant Germplasm Innovation and Utilization Institute of Environmental Horticulture, Guangdong Academy of Agricultural Sciences Guangzhou China
5. Department of Genetics and Biochemistry Clemson University Clemson SC USA