1. Key Laboratory of Horticultural Plant Biology (Ministry of Education) Key Laboratory of Horticultural Crop Biology and Genetic Improvement (Central Region, Ministry of Agriculture) College of Horticulture and Forestry Sciences Huazhong Agricultural University Wuhan 430070 China
2. Laboratory of Molecular Biology Department of Plant Sciences Wageningen University & Research Droevendaalsesteeg 16708 PBWageningen the Netherlands
3. Institute for Bioscience and Biotechnology Research & Department of Plant Sciences and Landscape Architecture University of Maryland College Park Rockville MD 20850 USA