1. Institute of Livestock and Grassland Science; National Agriculture and Food Research Organization; Tsukuba Ibaraki Japan
2. Kyushu Okinawa Agricultural Research Center; National Agriculture and Food Research Organization; Koshi Kumamoto Japan
3. Agricultural Research Center for Hokkaido Region; National Agriculture and Food Research Organization; Sapporo Hokkaido Japan
4. Institute of Environmental Management Technology; National Institute of Advanced Industrial Science and Technology; Tsukuba Ibaraki Japan
5. Department of Biological Sciences; Chuo University; Tokyo Japan