1. Institute of Evolutionary Biology, School of Biological Sciences; University of Edinburgh; Edinburgh United Kingdom
2. Division of Evolution and Ecology, Research School of Biology; The Australian National University; Canberra ACT Australia
3. Current Address: Nicolas Margraf, Musée d'histoire naturelle de La Chaux-de-Fonds; Av. Léopold-Robert 63 CH-2300 La Chaux-de-Fonds Switzerland